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Membrane separating the external ear from the middle ear

In the anatomy of humans and various other tetrapods, the eardrum, also called the tympanic membrane or myringa, is a thin, cone shaped membrane that separates the external ear from the middle ear. Its function is to transmit sound from the air to the ossicles inside the middle ear, and thence to the oval window in the fluid filled cochlea. The ear thereby converts and amplifies vibration in the air to vibration in cochlear fluid. The malleus bone bridges the gap between the eardrum and the other ossicles. Rupture or perforation of the eardrum can lead to conductive hearing loss. Collapse or retraction of the eardrum can cause conductive hearing loss or cholesteatoma.

The tympanic membrane is oriented obliquely in the anteroposterior, mediolateral, and superoinferior planes. Consequently, its superoposterior end lies lateral to its anteroinferior end. Anatomically, it relates superiorly to the middle cranial fossa, posteriorly to the ossicles and facial nerve, inferiorly to the parotid gland, and anteriorly to the temporomandibular joint.

The eardrum is divided into two general regions: the pars flaccida and the pars tensa. The relatively fragile pars flaccida lies above the lateral process of the malleus between the Notch of Rivinus and the anterior and posterior malleal folds. Consisting of two layers and appearing slightly pinkish in hue, it is associated with Eustachian tube dysfunction and cholesteatomas. The larger pars tensa consists of three layers: skin, fibrous tissue, and mucosa. Its thick periphery forms a fibrocartilaginous ring called the annulus tympanicus or Gerlach's ligament. while the central umbo tents inward at the level of the tip of malleus. The middle fibrous layer, containing radial, circular, and parabolic fibers, encloses the handle of malleus. Though comparatively robust, the pars tensa is the region more commonly associated with perforations.

The manubrium (Latin for "handle") of the malleus is firmly attached to the medial surface of the membrane as far as its center, drawing it toward the tympanic cavity. The lateral surface of the membrane is thus concave. The most depressed aspect of this concavity is termed the umbo (Latin for "shield boss").

Sensation of the outer surface of the tympanic membrane is supplied mainly by the auriculotemporal nerve, a branch of the mandibular nerve (cranial nerve V3), with contributions from the auricular branch of the vagus nerve (cranial nerve X), the facial nerve (cranial nerve VII), and possibly the glossopharyngeal nerve (cranial nerve IX). The inner surface of the tympanic membrane is innervated by the glossopharyngeal nerve.

When the eardrum is illuminated during a medical examination, a cone of light radiates from the tip of the malleus to the periphery in the anteroinferior quadrant, this is what is known clinically as 5 o'clock.

Unintentional perforation (rupture) has been described in blast injuries and air travel, typically in patients experiencing upper respiratory congestion or general Eustachian tube dysfunction that prevents equalization of pressure in the middle ear. It is also known to occur in swimming, diving (including scuba diving), and martial arts. Patients with tympanic membrane rupture may experience bleeding, tinnitus, hearing loss, or disequilibrium (vertigo). However, they rarely require medical intervention, as between 80 and 95 percent of ruptures recover completely within two to four weeks. The prognosis becomes more guarded as the force of injury increases.

The Bajau people of the Pacific intentionally rupture their eardrums at an early age to facilitate diving and hunting at sea. Many older Bajau therefore have difficulties hearing.