Введение
Вымерший род птицеподобных динозавров, древнее животное.
the ancient animal
Archaeopteryx (,/ɑːr//k//iː/'/ɒ//p//t//ər//ᵻ//k//s/; ), sometimes referred to by its German name, "Urvogel" ( Primeval Bird) is a genus of bird like dinosaurs. The name derives from the ancient Greek (archaīos), meaning "ancient", and (ptéryx), meaning "feather" or "wing". Between the late 19th century and the early 21st century, Archaeopteryx was generally accepted by palaeontologists and popular reference books as the oldest known bird (member of the group Avialae). Older potential avialans have since been identified, including Anchiornis, Xiaotingia, and Aurornis. Archaeopteryx lived in the Late Jurassic around 150 million years ago, in what is now southern Germany, during a time when Europe was an archipelago of islands in a shallow warm tropical sea, much closer to the equator than it is now. Similar in size to a Eurasian magpie, with the largest individuals possibly attaining the size of a raven, Most of these twelve fossils include impressions of feathers. Because these feathers are of an advanced form (flight feathers), these fossils are evidence that the evolution of feathers began before the Late Jurassic. It is currently located at the Natural History Museum of Berlin. Though it was the initial holotype, there were indications that it might not have been from the same animal as the body fossils. This conclusion was challenged in 2020 as being unlikely; the feather was identified on the basis of morphology as most likely having been an upper major primary covert feather.) to the Natural History Museum in London, where it remains. The Haarlem Specimen (TM 6428/29, also known as the Teylers Specimen) was discovered in 1855 near italics=unset, Germany, and described as a Pterodactylus crassipes in 1857 by Meyer. It was reclassified in 1970 by John Ostrom and is currently located at the Teylers Museum in Haarlem, the Netherlands. It was the very first specimen found, but was incorrectly classified at the time. It is also one of the least complete specimens, consisting mostly of limb bones, isolated cervical vertebrae, and ribs. In 2017 it was named as a separate genus Ostromia, considered more closely related to Anchiornis from China. The Eichstätt Specimen (JM 2257) was discovered in 1951 near Workerszell, Germany, and described by Peter Wellnhofer in 1974. Currently located at the Jura Museum in Eichstätt, Germany, it is the smallest known specimen and has the second best head. It is possibly a separate genus (Jurapteryx recurva) or species (A. recurva). The Solnhofen Specimen (unnumbered specimen) was discovered in the 1970s near Eichstätt, Germany, and described in 1988 by Wellnhofer. Currently located at the Bürgermeister Müller Museum in Solnhofen, it originally was classified as Compsognathus by an amateur collector, the same mayor Friedrich Müller after which the museum is named. It is the largest specimen known and may belong to a separate genus and species, Wellnhoferia grandis. It is missing only portions of the neck, tail, backbone, and head. The Munich Specimen (BSP 1999 I 50, formerly known as the Solenhofer Aktien Verein Specimen) was discovered on 3 August 1992 near Langenaltheim and described in 1993 by Wellnhofer. It is currently located at the Paläontologisches Museum München in Munich, to which it was sold in 1999 for 1.9 million Deutschmark. What was initially believed to be a bony sternum turned out to be part of the coracoid, but more recent studies suggest it belongs to A. siemensii. After a lengthy period in a closed private collection, it was moved to the Museum of Evolution at Knuthenborg Safaripark (Denmark) in 2022, where it has since been on display and also been made available for researchers. Another fragmentary fossil was found in 2000. It is in private possession and, since 2004, on loan to the Bürgermeister Müller Museum in Solnhofen, so it is called the Bürgermeister Müller Specimen; the institute itself officially refers to it as the "Exemplar of the families Ottman & Steil, Solnhofen". As the fragment represents the remains of a single wing of Archaeopteryx, it is colloquially known as "chicken wing". Long in a private collection in Switzerland, the Thermopolis Specimen (WDC CSG 100) was discovered in Bavaria and described in 2005 by Mayr, Pohl, and Peters. Donated to the Wyoming Dinosaur Center in Thermopolis, Wyoming, it has the best preserved head and feet; most of the neck and the lower jaw have not been preserved. The "Thermopolis" specimen was described on 2 December 2005 Science journal article as "A well preserved Archaeopteryx specimen with theropod features"; it shows that Archaeopteryx lacked a reversed toe—a universal feature of birds—limiting its ability to perch on branches and implying a terrestrial or trunk climbing lifestyle. Palaeontologists of the Ludwig Maximilian University of Munich studied the specimen, which revealed previously unknown features of the plumage, such as feathers on both the upper and lower legs and metatarsus, and the only preserved tail tip. A twelfth specimen had been discovered by an amateur collector in 2010 at the Schamhaupten quarry, but the finding was only announced in February 2014. It was scientifically described in 2018. It represents a complete and mostly articulated skeleton with skull. It is the only specimen lacking preserved feathers. It is from the Painten Formation and somewhat older than the other specimens. The existence of a thirteenth specimen (the Chicago specimen) was announced in 2024 by the Field Museum in Chicago, US. One of two specimens in an institution outside Europe, the specimen was originally identified in a private collection in Switzerland, and had been acquired by these collectors in 1990, prior to Germany's 2015 ban on exporting Archaeopteryx specimens. The specimen was acquired by the Field Museum in 2022, and went on public display in 2024 following two years of preparation. The specimen is to be studied by famed paleornithologist Jingmai O'Connor.
Археоптерикс (,/ɑːr//k//iː/'/ɒ//p//t//ər////k//s/;), иногда называемый его немецким названием "Urvogel" (Первоптица) — род птицеподобных динозавров. Название происходит от древнегреческого ἄρχαιος (archaīos), означающего «древний», и πτέρυξ (ptéryx), означающего «перо» или «крыло». С конца XIX века до начала XXI века археоптерикс обычно признавался палеонтологами и популярными справочниками как самая древняя известная птица (представитель группы Avialae). Впоследствии были идентифицированы более древние потенциальные авиаланы, включая Anchiornis, Xiaotingia и Aurornis. Археоптерикс жил в поздней юрской периодe около 150 миллионов лет назад на территории современной южной Германии, во времена, когда Европа представляла собой архипелаг островов в мелководном теплом тропическом море, расположенном гораздо ближе к экватору, чем сейчас. Большинство из этих двенадцати окаменелостей содержат отпечатки перьев. Поскольку эти перья имеют продвинутую форму (маховые перья), эти окаменелости свидетельствуют о том, что эволюция перьев началась до поздней юры. В настоящее время находится в Музее естественной истории в Берлине. Хотя изначально это был голотип, имелись предположения, что он мог принадлежать не тому же животному, что и образцы тела. Это заключение было оспорено в 2020 году как маловероятное; перо было идентифицировано по морфологии как, скорее всего, верхнее маховое покровное перо. Образец был передан в Музей естественной истории в Лондоне, где он и находится по сей день.
the ancient animal
Archaeopteryx (,/ɑːr//k//iː/'/ɒ//p//t//ər//ᵻ//k//s/; ), sometimes referred to by its German name, "Urvogel" ( Primeval Bird) is a genus of bird like dinosaurs. The name derives from the ancient Greek (archaīos), meaning "ancient", and (ptéryx), meaning "feather" or "wing". Between the late 19th century and the early 21st century, Archaeopteryx was generally accepted by palaeontologists and popular reference books as the oldest known bird (member of the group Avialae). Older potential avialans have since been identified, including Anchiornis, Xiaotingia, and Aurornis. Archaeopteryx lived in the Late Jurassic around 150 million years ago, in what is now southern Germany, during a time when Europe was an archipelago of islands in a shallow warm tropical sea, much closer to the equator than it is now. Similar in size to a Eurasian magpie, with the largest individuals possibly attaining the size of a raven, Most of these twelve fossils include impressions of feathers. Because these feathers are of an advanced form (flight feathers), these fossils are evidence that the evolution of feathers began before the Late Jurassic. It is currently located at the Natural History Museum of Berlin. Though it was the initial holotype, there were indications that it might not have been from the same animal as the body fossils. This conclusion was challenged in 2020 as being unlikely; the feather was identified on the basis of morphology as most likely having been an upper major primary covert feather.) to the Natural History Museum in London, where it remains. The Haarlem Specimen (TM 6428/29, also known as the Teylers Specimen) was discovered in 1855 near italics=unset, Germany, and described as a Pterodactylus crassipes in 1857 by Meyer. It was reclassified in 1970 by John Ostrom and is currently located at the Teylers Museum in Haarlem, the Netherlands. It was the very first specimen found, but was incorrectly classified at the time. It is also one of the least complete specimens, consisting mostly of limb bones, isolated cervical vertebrae, and ribs. In 2017 it was named as a separate genus Ostromia, considered more closely related to Anchiornis from China. The Eichstätt Specimen (JM 2257) was discovered in 1951 near Workerszell, Germany, and described by Peter Wellnhofer in 1974. Currently located at the Jura Museum in Eichstätt, Germany, it is the smallest known specimen and has the second best head. It is possibly a separate genus (Jurapteryx recurva) or species (A. recurva). The Solnhofen Specimen (unnumbered specimen) was discovered in the 1970s near Eichstätt, Germany, and described in 1988 by Wellnhofer. Currently located at the Bürgermeister Müller Museum in Solnhofen, it originally was classified as Compsognathus by an amateur collector, the same mayor Friedrich Müller after which the museum is named. It is the largest specimen known and may belong to a separate genus and species, Wellnhoferia grandis. It is missing only portions of the neck, tail, backbone, and head. The Munich Specimen (BSP 1999 I 50, formerly known as the Solenhofer Aktien Verein Specimen) was discovered on 3 August 1992 near Langenaltheim and described in 1993 by Wellnhofer. It is currently located at the Paläontologisches Museum München in Munich, to which it was sold in 1999 for 1.9 million Deutschmark. What was initially believed to be a bony sternum turned out to be part of the coracoid, but more recent studies suggest it belongs to A. siemensii. After a lengthy period in a closed private collection, it was moved to the Museum of Evolution at Knuthenborg Safaripark (Denmark) in 2022, where it has since been on display and also been made available for researchers. Another fragmentary fossil was found in 2000. It is in private possession and, since 2004, on loan to the Bürgermeister Müller Museum in Solnhofen, so it is called the Bürgermeister Müller Specimen; the institute itself officially refers to it as the "Exemplar of the families Ottman & Steil, Solnhofen". As the fragment represents the remains of a single wing of Archaeopteryx, it is colloquially known as "chicken wing". Long in a private collection in Switzerland, the Thermopolis Specimen (WDC CSG 100) was discovered in Bavaria and described in 2005 by Mayr, Pohl, and Peters. Donated to the Wyoming Dinosaur Center in Thermopolis, Wyoming, it has the best preserved head and feet; most of the neck and the lower jaw have not been preserved. The "Thermopolis" specimen was described on 2 December 2005 Science journal article as "A well preserved Archaeopteryx specimen with theropod features"; it shows that Archaeopteryx lacked a reversed toe—a universal feature of birds—limiting its ability to perch on branches and implying a terrestrial or trunk climbing lifestyle. Palaeontologists of the Ludwig Maximilian University of Munich studied the specimen, which revealed previously unknown features of the plumage, such as feathers on both the upper and lower legs and metatarsus, and the only preserved tail tip. A twelfth specimen had been discovered by an amateur collector in 2010 at the Schamhaupten quarry, but the finding was only announced in February 2014. It was scientifically described in 2018. It represents a complete and mostly articulated skeleton with skull. It is the only specimen lacking preserved feathers. It is from the Painten Formation and somewhat older than the other specimens. The existence of a thirteenth specimen (the Chicago specimen) was announced in 2024 by the Field Museum in Chicago, US. One of two specimens in an institution outside Europe, the specimen was originally identified in a private collection in Switzerland, and had been acquired by these collectors in 1990, prior to Germany's 2015 ban on exporting Archaeopteryx specimens. The specimen was acquired by the Field Museum in 2022, and went on public display in 2024 following two years of preparation. The specimen is to be studied by famed paleornithologist Jingmai O'Connor.
Хаарлемский образец (TM 6428/29, также известный как образец Тейлера) был обнаружен в 1855 году недалеко от Германии и описан Мейером в 1857 году как Pterodactylus crassipes. В 1970 году он был переклассифицирован Джоном Остромом и в настоящее время находится в музее Тейлера в Харлеме, Нидерланды. Это был первый найденный образец, но в то время он был неправильно классифицирован. Это также один из наименее полных образцов, состоящий в основном из костей конечностей, изолированных шейных позвонков и ребер. В 2017 году он был отнесен к отдельному роду Ostromia, который считается более близким к Anchiornis из Китая.
the ancient animal
Archaeopteryx (,/ɑːr//k//iː/'/ɒ//p//t//ər//ᵻ//k//s/; ), sometimes referred to by its German name, "Urvogel" ( Primeval Bird) is a genus of bird like dinosaurs. The name derives from the ancient Greek (archaīos), meaning "ancient", and (ptéryx), meaning "feather" or "wing". Between the late 19th century and the early 21st century, Archaeopteryx was generally accepted by palaeontologists and popular reference books as the oldest known bird (member of the group Avialae). Older potential avialans have since been identified, including Anchiornis, Xiaotingia, and Aurornis. Archaeopteryx lived in the Late Jurassic around 150 million years ago, in what is now southern Germany, during a time when Europe was an archipelago of islands in a shallow warm tropical sea, much closer to the equator than it is now. Similar in size to a Eurasian magpie, with the largest individuals possibly attaining the size of a raven, Most of these twelve fossils include impressions of feathers. Because these feathers are of an advanced form (flight feathers), these fossils are evidence that the evolution of feathers began before the Late Jurassic. It is currently located at the Natural History Museum of Berlin. Though it was the initial holotype, there were indications that it might not have been from the same animal as the body fossils. This conclusion was challenged in 2020 as being unlikely; the feather was identified on the basis of morphology as most likely having been an upper major primary covert feather.) to the Natural History Museum in London, where it remains. The Haarlem Specimen (TM 6428/29, also known as the Teylers Specimen) was discovered in 1855 near italics=unset, Germany, and described as a Pterodactylus crassipes in 1857 by Meyer. It was reclassified in 1970 by John Ostrom and is currently located at the Teylers Museum in Haarlem, the Netherlands. It was the very first specimen found, but was incorrectly classified at the time. It is also one of the least complete specimens, consisting mostly of limb bones, isolated cervical vertebrae, and ribs. In 2017 it was named as a separate genus Ostromia, considered more closely related to Anchiornis from China. The Eichstätt Specimen (JM 2257) was discovered in 1951 near Workerszell, Germany, and described by Peter Wellnhofer in 1974. Currently located at the Jura Museum in Eichstätt, Germany, it is the smallest known specimen and has the second best head. It is possibly a separate genus (Jurapteryx recurva) or species (A. recurva). The Solnhofen Specimen (unnumbered specimen) was discovered in the 1970s near Eichstätt, Germany, and described in 1988 by Wellnhofer. Currently located at the Bürgermeister Müller Museum in Solnhofen, it originally was classified as Compsognathus by an amateur collector, the same mayor Friedrich Müller after which the museum is named. It is the largest specimen known and may belong to a separate genus and species, Wellnhoferia grandis. It is missing only portions of the neck, tail, backbone, and head. The Munich Specimen (BSP 1999 I 50, formerly known as the Solenhofer Aktien Verein Specimen) was discovered on 3 August 1992 near Langenaltheim and described in 1993 by Wellnhofer. It is currently located at the Paläontologisches Museum München in Munich, to which it was sold in 1999 for 1.9 million Deutschmark. What was initially believed to be a bony sternum turned out to be part of the coracoid, but more recent studies suggest it belongs to A. siemensii. After a lengthy period in a closed private collection, it was moved to the Museum of Evolution at Knuthenborg Safaripark (Denmark) in 2022, where it has since been on display and also been made available for researchers. Another fragmentary fossil was found in 2000. It is in private possession and, since 2004, on loan to the Bürgermeister Müller Museum in Solnhofen, so it is called the Bürgermeister Müller Specimen; the institute itself officially refers to it as the "Exemplar of the families Ottman & Steil, Solnhofen". As the fragment represents the remains of a single wing of Archaeopteryx, it is colloquially known as "chicken wing". Long in a private collection in Switzerland, the Thermopolis Specimen (WDC CSG 100) was discovered in Bavaria and described in 2005 by Mayr, Pohl, and Peters. Donated to the Wyoming Dinosaur Center in Thermopolis, Wyoming, it has the best preserved head and feet; most of the neck and the lower jaw have not been preserved. The "Thermopolis" specimen was described on 2 December 2005 Science journal article as "A well preserved Archaeopteryx specimen with theropod features"; it shows that Archaeopteryx lacked a reversed toe—a universal feature of birds—limiting its ability to perch on branches and implying a terrestrial or trunk climbing lifestyle. Palaeontologists of the Ludwig Maximilian University of Munich studied the specimen, which revealed previously unknown features of the plumage, such as feathers on both the upper and lower legs and metatarsus, and the only preserved tail tip. A twelfth specimen had been discovered by an amateur collector in 2010 at the Schamhaupten quarry, but the finding was only announced in February 2014. It was scientifically described in 2018. It represents a complete and mostly articulated skeleton with skull. It is the only specimen lacking preserved feathers. It is from the Painten Formation and somewhat older than the other specimens. The existence of a thirteenth specimen (the Chicago specimen) was announced in 2024 by the Field Museum in Chicago, US. One of two specimens in an institution outside Europe, the specimen was originally identified in a private collection in Switzerland, and had been acquired by these collectors in 1990, prior to Germany's 2015 ban on exporting Archaeopteryx specimens. The specimen was acquired by the Field Museum in 2022, and went on public display in 2024 following two years of preparation. The specimen is to be studied by famed paleornithologist Jingmai O'Connor.
Образец из Эйхштетта (JM 2257) был обнаружен в 1951 году недалеко от города Веркерсцелль, Германия, и описан Питером Велленхофером в 1974 году. В настоящее время он находится в Музее Юры в Эйхштетте, Германия, и является самым маленьким известным образцом, с второй по величине головой. Возможно, это отдельный род (Jurapteryx recurva) или вид (A. recurva).
the ancient animal
Archaeopteryx (,/ɑːr//k//iː/'/ɒ//p//t//ər//ᵻ//k//s/; ), sometimes referred to by its German name, "Urvogel" ( Primeval Bird) is a genus of bird like dinosaurs. The name derives from the ancient Greek (archaīos), meaning "ancient", and (ptéryx), meaning "feather" or "wing". Between the late 19th century and the early 21st century, Archaeopteryx was generally accepted by palaeontologists and popular reference books as the oldest known bird (member of the group Avialae). Older potential avialans have since been identified, including Anchiornis, Xiaotingia, and Aurornis. Archaeopteryx lived in the Late Jurassic around 150 million years ago, in what is now southern Germany, during a time when Europe was an archipelago of islands in a shallow warm tropical sea, much closer to the equator than it is now. Similar in size to a Eurasian magpie, with the largest individuals possibly attaining the size of a raven, Most of these twelve fossils include impressions of feathers. Because these feathers are of an advanced form (flight feathers), these fossils are evidence that the evolution of feathers began before the Late Jurassic. It is currently located at the Natural History Museum of Berlin. Though it was the initial holotype, there were indications that it might not have been from the same animal as the body fossils. This conclusion was challenged in 2020 as being unlikely; the feather was identified on the basis of morphology as most likely having been an upper major primary covert feather.) to the Natural History Museum in London, where it remains. The Haarlem Specimen (TM 6428/29, also known as the Teylers Specimen) was discovered in 1855 near italics=unset, Germany, and described as a Pterodactylus crassipes in 1857 by Meyer. It was reclassified in 1970 by John Ostrom and is currently located at the Teylers Museum in Haarlem, the Netherlands. It was the very first specimen found, but was incorrectly classified at the time. It is also one of the least complete specimens, consisting mostly of limb bones, isolated cervical vertebrae, and ribs. In 2017 it was named as a separate genus Ostromia, considered more closely related to Anchiornis from China. The Eichstätt Specimen (JM 2257) was discovered in 1951 near Workerszell, Germany, and described by Peter Wellnhofer in 1974. Currently located at the Jura Museum in Eichstätt, Germany, it is the smallest known specimen and has the second best head. It is possibly a separate genus (Jurapteryx recurva) or species (A. recurva). The Solnhofen Specimen (unnumbered specimen) was discovered in the 1970s near Eichstätt, Germany, and described in 1988 by Wellnhofer. Currently located at the Bürgermeister Müller Museum in Solnhofen, it originally was classified as Compsognathus by an amateur collector, the same mayor Friedrich Müller after which the museum is named. It is the largest specimen known and may belong to a separate genus and species, Wellnhoferia grandis. It is missing only portions of the neck, tail, backbone, and head. The Munich Specimen (BSP 1999 I 50, formerly known as the Solenhofer Aktien Verein Specimen) was discovered on 3 August 1992 near Langenaltheim and described in 1993 by Wellnhofer. It is currently located at the Paläontologisches Museum München in Munich, to which it was sold in 1999 for 1.9 million Deutschmark. What was initially believed to be a bony sternum turned out to be part of the coracoid, but more recent studies suggest it belongs to A. siemensii. After a lengthy period in a closed private collection, it was moved to the Museum of Evolution at Knuthenborg Safaripark (Denmark) in 2022, where it has since been on display and also been made available for researchers. Another fragmentary fossil was found in 2000. It is in private possession and, since 2004, on loan to the Bürgermeister Müller Museum in Solnhofen, so it is called the Bürgermeister Müller Specimen; the institute itself officially refers to it as the "Exemplar of the families Ottman & Steil, Solnhofen". As the fragment represents the remains of a single wing of Archaeopteryx, it is colloquially known as "chicken wing". Long in a private collection in Switzerland, the Thermopolis Specimen (WDC CSG 100) was discovered in Bavaria and described in 2005 by Mayr, Pohl, and Peters. Donated to the Wyoming Dinosaur Center in Thermopolis, Wyoming, it has the best preserved head and feet; most of the neck and the lower jaw have not been preserved. The "Thermopolis" specimen was described on 2 December 2005 Science journal article as "A well preserved Archaeopteryx specimen with theropod features"; it shows that Archaeopteryx lacked a reversed toe—a universal feature of birds—limiting its ability to perch on branches and implying a terrestrial or trunk climbing lifestyle. Palaeontologists of the Ludwig Maximilian University of Munich studied the specimen, which revealed previously unknown features of the plumage, such as feathers on both the upper and lower legs and metatarsus, and the only preserved tail tip. A twelfth specimen had been discovered by an amateur collector in 2010 at the Schamhaupten quarry, but the finding was only announced in February 2014. It was scientifically described in 2018. It represents a complete and mostly articulated skeleton with skull. It is the only specimen lacking preserved feathers. It is from the Painten Formation and somewhat older than the other specimens. The existence of a thirteenth specimen (the Chicago specimen) was announced in 2024 by the Field Museum in Chicago, US. One of two specimens in an institution outside Europe, the specimen was originally identified in a private collection in Switzerland, and had been acquired by these collectors in 1990, prior to Germany's 2015 ban on exporting Archaeopteryx specimens. The specimen was acquired by the Field Museum in 2022, and went on public display in 2024 following two years of preparation. The specimen is to be studied by famed paleornithologist Jingmai O'Connor.
Образец из Солнхофена (непронумерованный образец) был обнаружен в 1970-х годах недалеко от Эйхштетта, Германия, и описан Велленхофером в 1988 году. В настоящее время он находится в музее Бюргермейстера Мюллера в Солнхофене, где изначально был классифицирован как Compsognathus любительским коллекционером, тем же мэром Фридрихом Мюллером, в честь которого назван музей. Это самый большой известный образец, который может принадлежать к отдельному роду и виду, Wellnhoferia grandis. От него отсутствуют только части шеи, хвоста, позвоночника и головы.
the ancient animal
Archaeopteryx (,/ɑːr//k//iː/'/ɒ//p//t//ər//ᵻ//k//s/; ), sometimes referred to by its German name, "Urvogel" ( Primeval Bird) is a genus of bird like dinosaurs. The name derives from the ancient Greek (archaīos), meaning "ancient", and (ptéryx), meaning "feather" or "wing". Between the late 19th century and the early 21st century, Archaeopteryx was generally accepted by palaeontologists and popular reference books as the oldest known bird (member of the group Avialae). Older potential avialans have since been identified, including Anchiornis, Xiaotingia, and Aurornis. Archaeopteryx lived in the Late Jurassic around 150 million years ago, in what is now southern Germany, during a time when Europe was an archipelago of islands in a shallow warm tropical sea, much closer to the equator than it is now. Similar in size to a Eurasian magpie, with the largest individuals possibly attaining the size of a raven, Most of these twelve fossils include impressions of feathers. Because these feathers are of an advanced form (flight feathers), these fossils are evidence that the evolution of feathers began before the Late Jurassic. It is currently located at the Natural History Museum of Berlin. Though it was the initial holotype, there were indications that it might not have been from the same animal as the body fossils. This conclusion was challenged in 2020 as being unlikely; the feather was identified on the basis of morphology as most likely having been an upper major primary covert feather.) to the Natural History Museum in London, where it remains. The Haarlem Specimen (TM 6428/29, also known as the Teylers Specimen) was discovered in 1855 near italics=unset, Germany, and described as a Pterodactylus crassipes in 1857 by Meyer. It was reclassified in 1970 by John Ostrom and is currently located at the Teylers Museum in Haarlem, the Netherlands. It was the very first specimen found, but was incorrectly classified at the time. It is also one of the least complete specimens, consisting mostly of limb bones, isolated cervical vertebrae, and ribs. In 2017 it was named as a separate genus Ostromia, considered more closely related to Anchiornis from China. The Eichstätt Specimen (JM 2257) was discovered in 1951 near Workerszell, Germany, and described by Peter Wellnhofer in 1974. Currently located at the Jura Museum in Eichstätt, Germany, it is the smallest known specimen and has the second best head. It is possibly a separate genus (Jurapteryx recurva) or species (A. recurva). The Solnhofen Specimen (unnumbered specimen) was discovered in the 1970s near Eichstätt, Germany, and described in 1988 by Wellnhofer. Currently located at the Bürgermeister Müller Museum in Solnhofen, it originally was classified as Compsognathus by an amateur collector, the same mayor Friedrich Müller after which the museum is named. It is the largest specimen known and may belong to a separate genus and species, Wellnhoferia grandis. It is missing only portions of the neck, tail, backbone, and head. The Munich Specimen (BSP 1999 I 50, formerly known as the Solenhofer Aktien Verein Specimen) was discovered on 3 August 1992 near Langenaltheim and described in 1993 by Wellnhofer. It is currently located at the Paläontologisches Museum München in Munich, to which it was sold in 1999 for 1.9 million Deutschmark. What was initially believed to be a bony sternum turned out to be part of the coracoid, but more recent studies suggest it belongs to A. siemensii. After a lengthy period in a closed private collection, it was moved to the Museum of Evolution at Knuthenborg Safaripark (Denmark) in 2022, where it has since been on display and also been made available for researchers. Another fragmentary fossil was found in 2000. It is in private possession and, since 2004, on loan to the Bürgermeister Müller Museum in Solnhofen, so it is called the Bürgermeister Müller Specimen; the institute itself officially refers to it as the "Exemplar of the families Ottman & Steil, Solnhofen". As the fragment represents the remains of a single wing of Archaeopteryx, it is colloquially known as "chicken wing". Long in a private collection in Switzerland, the Thermopolis Specimen (WDC CSG 100) was discovered in Bavaria and described in 2005 by Mayr, Pohl, and Peters. Donated to the Wyoming Dinosaur Center in Thermopolis, Wyoming, it has the best preserved head and feet; most of the neck and the lower jaw have not been preserved. The "Thermopolis" specimen was described on 2 December 2005 Science journal article as "A well preserved Archaeopteryx specimen with theropod features"; it shows that Archaeopteryx lacked a reversed toe—a universal feature of birds—limiting its ability to perch on branches and implying a terrestrial or trunk climbing lifestyle. Palaeontologists of the Ludwig Maximilian University of Munich studied the specimen, which revealed previously unknown features of the plumage, such as feathers on both the upper and lower legs and metatarsus, and the only preserved tail tip. A twelfth specimen had been discovered by an amateur collector in 2010 at the Schamhaupten quarry, but the finding was only announced in February 2014. It was scientifically described in 2018. It represents a complete and mostly articulated skeleton with skull. It is the only specimen lacking preserved feathers. It is from the Painten Formation and somewhat older than the other specimens. The existence of a thirteenth specimen (the Chicago specimen) was announced in 2024 by the Field Museum in Chicago, US. One of two specimens in an institution outside Europe, the specimen was originally identified in a private collection in Switzerland, and had been acquired by these collectors in 1990, prior to Germany's 2015 ban on exporting Archaeopteryx specimens. The specimen was acquired by the Field Museum in 2022, and went on public display in 2024 following two years of preparation. The specimen is to be studied by famed paleornithologist Jingmai O'Connor.
Мюнхенский образец (BSP 1999 I 50, ранее известный как образец Solenhofer Aktien Verein) был обнаружен 3 августа 1992 года недалеко от Лангенальтейма и описан Велленхофером в 1993 году. В настоящее время он находится в Палеонтологическом музее в Мюнхене, которому был продан в 1999 году за 1,9 миллиона немецких марок. То, что первоначально считалось костной грудиной, оказалось частью коракоида, но более поздние исследования показывают, что оно принадлежит A. siemensii. После длительного периода в закрытой частной коллекции в 2022 году он был передан в Музей эволюции в Сафари-парке Кнутенборг (Дания), где с тех пор он выставлен и доступен исследователям.
the ancient animal
Archaeopteryx (,/ɑːr//k//iː/'/ɒ//p//t//ər//ᵻ//k//s/; ), sometimes referred to by its German name, "Urvogel" ( Primeval Bird) is a genus of bird like dinosaurs. The name derives from the ancient Greek (archaīos), meaning "ancient", and (ptéryx), meaning "feather" or "wing". Between the late 19th century and the early 21st century, Archaeopteryx was generally accepted by palaeontologists and popular reference books as the oldest known bird (member of the group Avialae). Older potential avialans have since been identified, including Anchiornis, Xiaotingia, and Aurornis. Archaeopteryx lived in the Late Jurassic around 150 million years ago, in what is now southern Germany, during a time when Europe was an archipelago of islands in a shallow warm tropical sea, much closer to the equator than it is now. Similar in size to a Eurasian magpie, with the largest individuals possibly attaining the size of a raven, Most of these twelve fossils include impressions of feathers. Because these feathers are of an advanced form (flight feathers), these fossils are evidence that the evolution of feathers began before the Late Jurassic. It is currently located at the Natural History Museum of Berlin. Though it was the initial holotype, there were indications that it might not have been from the same animal as the body fossils. This conclusion was challenged in 2020 as being unlikely; the feather was identified on the basis of morphology as most likely having been an upper major primary covert feather.) to the Natural History Museum in London, where it remains. The Haarlem Specimen (TM 6428/29, also known as the Teylers Specimen) was discovered in 1855 near italics=unset, Germany, and described as a Pterodactylus crassipes in 1857 by Meyer. It was reclassified in 1970 by John Ostrom and is currently located at the Teylers Museum in Haarlem, the Netherlands. It was the very first specimen found, but was incorrectly classified at the time. It is also one of the least complete specimens, consisting mostly of limb bones, isolated cervical vertebrae, and ribs. In 2017 it was named as a separate genus Ostromia, considered more closely related to Anchiornis from China. The Eichstätt Specimen (JM 2257) was discovered in 1951 near Workerszell, Germany, and described by Peter Wellnhofer in 1974. Currently located at the Jura Museum in Eichstätt, Germany, it is the smallest known specimen and has the second best head. It is possibly a separate genus (Jurapteryx recurva) or species (A. recurva). The Solnhofen Specimen (unnumbered specimen) was discovered in the 1970s near Eichstätt, Germany, and described in 1988 by Wellnhofer. Currently located at the Bürgermeister Müller Museum in Solnhofen, it originally was classified as Compsognathus by an amateur collector, the same mayor Friedrich Müller after which the museum is named. It is the largest specimen known and may belong to a separate genus and species, Wellnhoferia grandis. It is missing only portions of the neck, tail, backbone, and head. The Munich Specimen (BSP 1999 I 50, formerly known as the Solenhofer Aktien Verein Specimen) was discovered on 3 August 1992 near Langenaltheim and described in 1993 by Wellnhofer. It is currently located at the Paläontologisches Museum München in Munich, to which it was sold in 1999 for 1.9 million Deutschmark. What was initially believed to be a bony sternum turned out to be part of the coracoid, but more recent studies suggest it belongs to A. siemensii. After a lengthy period in a closed private collection, it was moved to the Museum of Evolution at Knuthenborg Safaripark (Denmark) in 2022, where it has since been on display and also been made available for researchers. Another fragmentary fossil was found in 2000. It is in private possession and, since 2004, on loan to the Bürgermeister Müller Museum in Solnhofen, so it is called the Bürgermeister Müller Specimen; the institute itself officially refers to it as the "Exemplar of the families Ottman & Steil, Solnhofen". As the fragment represents the remains of a single wing of Archaeopteryx, it is colloquially known as "chicken wing". Long in a private collection in Switzerland, the Thermopolis Specimen (WDC CSG 100) was discovered in Bavaria and described in 2005 by Mayr, Pohl, and Peters. Donated to the Wyoming Dinosaur Center in Thermopolis, Wyoming, it has the best preserved head and feet; most of the neck and the lower jaw have not been preserved. The "Thermopolis" specimen was described on 2 December 2005 Science journal article as "A well preserved Archaeopteryx specimen with theropod features"; it shows that Archaeopteryx lacked a reversed toe—a universal feature of birds—limiting its ability to perch on branches and implying a terrestrial or trunk climbing lifestyle. Palaeontologists of the Ludwig Maximilian University of Munich studied the specimen, which revealed previously unknown features of the plumage, such as feathers on both the upper and lower legs and metatarsus, and the only preserved tail tip. A twelfth specimen had been discovered by an amateur collector in 2010 at the Schamhaupten quarry, but the finding was only announced in February 2014. It was scientifically described in 2018. It represents a complete and mostly articulated skeleton with skull. It is the only specimen lacking preserved feathers. It is from the Painten Formation and somewhat older than the other specimens. The existence of a thirteenth specimen (the Chicago specimen) was announced in 2024 by the Field Museum in Chicago, US. One of two specimens in an institution outside Europe, the specimen was originally identified in a private collection in Switzerland, and had been acquired by these collectors in 1990, prior to Germany's 2015 ban on exporting Archaeopteryx specimens. The specimen was acquired by the Field Museum in 2022, and went on public display in 2024 following two years of preparation. The specimen is to be studied by famed paleornithologist Jingmai O'Connor.
Еще один фрагментарный окаменелость был найден в 2000 году. Он находится в частном владении и с 2004 года находится в аренде у музея Бюргермейстера Мюллера в Солнхофене, поэтому его называют образцом Бюргермейстера Мюллера; сам институт официально называет его «образцом семейств Ottman & Steil, Solnhofen». Поскольку фрагмент представляет собой остатки одного крыла археоптерикса, он в разговорной речи известен как «куриное крыло».
the ancient animal
Archaeopteryx (,/ɑːr//k//iː/'/ɒ//p//t//ər//ᵻ//k//s/; ), sometimes referred to by its German name, "Urvogel" ( Primeval Bird) is a genus of bird like dinosaurs. The name derives from the ancient Greek (archaīos), meaning "ancient", and (ptéryx), meaning "feather" or "wing". Between the late 19th century and the early 21st century, Archaeopteryx was generally accepted by palaeontologists and popular reference books as the oldest known bird (member of the group Avialae). Older potential avialans have since been identified, including Anchiornis, Xiaotingia, and Aurornis. Archaeopteryx lived in the Late Jurassic around 150 million years ago, in what is now southern Germany, during a time when Europe was an archipelago of islands in a shallow warm tropical sea, much closer to the equator than it is now. Similar in size to a Eurasian magpie, with the largest individuals possibly attaining the size of a raven, Most of these twelve fossils include impressions of feathers. Because these feathers are of an advanced form (flight feathers), these fossils are evidence that the evolution of feathers began before the Late Jurassic. It is currently located at the Natural History Museum of Berlin. Though it was the initial holotype, there were indications that it might not have been from the same animal as the body fossils. This conclusion was challenged in 2020 as being unlikely; the feather was identified on the basis of morphology as most likely having been an upper major primary covert feather.) to the Natural History Museum in London, where it remains. The Haarlem Specimen (TM 6428/29, also known as the Teylers Specimen) was discovered in 1855 near italics=unset, Germany, and described as a Pterodactylus crassipes in 1857 by Meyer. It was reclassified in 1970 by John Ostrom and is currently located at the Teylers Museum in Haarlem, the Netherlands. It was the very first specimen found, but was incorrectly classified at the time. It is also one of the least complete specimens, consisting mostly of limb bones, isolated cervical vertebrae, and ribs. In 2017 it was named as a separate genus Ostromia, considered more closely related to Anchiornis from China. The Eichstätt Specimen (JM 2257) was discovered in 1951 near Workerszell, Germany, and described by Peter Wellnhofer in 1974. Currently located at the Jura Museum in Eichstätt, Germany, it is the smallest known specimen and has the second best head. It is possibly a separate genus (Jurapteryx recurva) or species (A. recurva). The Solnhofen Specimen (unnumbered specimen) was discovered in the 1970s near Eichstätt, Germany, and described in 1988 by Wellnhofer. Currently located at the Bürgermeister Müller Museum in Solnhofen, it originally was classified as Compsognathus by an amateur collector, the same mayor Friedrich Müller after which the museum is named. It is the largest specimen known and may belong to a separate genus and species, Wellnhoferia grandis. It is missing only portions of the neck, tail, backbone, and head. The Munich Specimen (BSP 1999 I 50, formerly known as the Solenhofer Aktien Verein Specimen) was discovered on 3 August 1992 near Langenaltheim and described in 1993 by Wellnhofer. It is currently located at the Paläontologisches Museum München in Munich, to which it was sold in 1999 for 1.9 million Deutschmark. What was initially believed to be a bony sternum turned out to be part of the coracoid, but more recent studies suggest it belongs to A. siemensii. After a lengthy period in a closed private collection, it was moved to the Museum of Evolution at Knuthenborg Safaripark (Denmark) in 2022, where it has since been on display and also been made available for researchers. Another fragmentary fossil was found in 2000. It is in private possession and, since 2004, on loan to the Bürgermeister Müller Museum in Solnhofen, so it is called the Bürgermeister Müller Specimen; the institute itself officially refers to it as the "Exemplar of the families Ottman & Steil, Solnhofen". As the fragment represents the remains of a single wing of Archaeopteryx, it is colloquially known as "chicken wing". Long in a private collection in Switzerland, the Thermopolis Specimen (WDC CSG 100) was discovered in Bavaria and described in 2005 by Mayr, Pohl, and Peters. Donated to the Wyoming Dinosaur Center in Thermopolis, Wyoming, it has the best preserved head and feet; most of the neck and the lower jaw have not been preserved. The "Thermopolis" specimen was described on 2 December 2005 Science journal article as "A well preserved Archaeopteryx specimen with theropod features"; it shows that Archaeopteryx lacked a reversed toe—a universal feature of birds—limiting its ability to perch on branches and implying a terrestrial or trunk climbing lifestyle. Palaeontologists of the Ludwig Maximilian University of Munich studied the specimen, which revealed previously unknown features of the plumage, such as feathers on both the upper and lower legs and metatarsus, and the only preserved tail tip. A twelfth specimen had been discovered by an amateur collector in 2010 at the Schamhaupten quarry, but the finding was only announced in February 2014. It was scientifically described in 2018. It represents a complete and mostly articulated skeleton with skull. It is the only specimen lacking preserved feathers. It is from the Painten Formation and somewhat older than the other specimens. The existence of a thirteenth specimen (the Chicago specimen) was announced in 2024 by the Field Museum in Chicago, US. One of two specimens in an institution outside Europe, the specimen was originally identified in a private collection in Switzerland, and had been acquired by these collectors in 1990, prior to Germany's 2015 ban on exporting Archaeopteryx specimens. The specimen was acquired by the Field Museum in 2022, and went on public display in 2024 following two years of preparation. The specimen is to be studied by famed paleornithologist Jingmai O'Connor.
Долгое время находившийся в частной коллекции в Швейцарии, образец Термополиса (WDC CSG 100) был обнаружен в Баварии и описан в 2005 году Майром, Полом и Петерсом. Пожертвованный в Центр динозавров Вайоминга в Термополисе, штат Вайоминг, он имеет наиболее хорошо сохранившуюся голову и ноги; большая часть шеи и нижней челюсти не сохранились. В статье журнала Science от 2 декабря 2005 года образец «Термополис» был описан как «Хорошо сохранившийся образец археоптерикса с чертами теропода»; в нем показано, что у археоптерикса отсутствовал обращенный назад палец — универсальная черта птиц — что ограничивало его способность садиться на ветки и предполагало наземный или древесный образ жизни. Палеонтологи из Людвиг-Максимилианского университета Мюнхена изучили образец, который выявил ранее неизвестные особенности оперения, такие как перья как на верхней, так и на нижней ногах и плюсне, а также единственный сохранившийся кончик хвоста.
the ancient animal
Archaeopteryx (,/ɑːr//k//iː/'/ɒ//p//t//ər//ᵻ//k//s/; ), sometimes referred to by its German name, "Urvogel" ( Primeval Bird) is a genus of bird like dinosaurs. The name derives from the ancient Greek (archaīos), meaning "ancient", and (ptéryx), meaning "feather" or "wing". Between the late 19th century and the early 21st century, Archaeopteryx was generally accepted by palaeontologists and popular reference books as the oldest known bird (member of the group Avialae). Older potential avialans have since been identified, including Anchiornis, Xiaotingia, and Aurornis. Archaeopteryx lived in the Late Jurassic around 150 million years ago, in what is now southern Germany, during a time when Europe was an archipelago of islands in a shallow warm tropical sea, much closer to the equator than it is now. Similar in size to a Eurasian magpie, with the largest individuals possibly attaining the size of a raven, Most of these twelve fossils include impressions of feathers. Because these feathers are of an advanced form (flight feathers), these fossils are evidence that the evolution of feathers began before the Late Jurassic. It is currently located at the Natural History Museum of Berlin. Though it was the initial holotype, there were indications that it might not have been from the same animal as the body fossils. This conclusion was challenged in 2020 as being unlikely; the feather was identified on the basis of morphology as most likely having been an upper major primary covert feather.) to the Natural History Museum in London, where it remains. The Haarlem Specimen (TM 6428/29, also known as the Teylers Specimen) was discovered in 1855 near italics=unset, Germany, and described as a Pterodactylus crassipes in 1857 by Meyer. It was reclassified in 1970 by John Ostrom and is currently located at the Teylers Museum in Haarlem, the Netherlands. It was the very first specimen found, but was incorrectly classified at the time. It is also one of the least complete specimens, consisting mostly of limb bones, isolated cervical vertebrae, and ribs. In 2017 it was named as a separate genus Ostromia, considered more closely related to Anchiornis from China. The Eichstätt Specimen (JM 2257) was discovered in 1951 near Workerszell, Germany, and described by Peter Wellnhofer in 1974. Currently located at the Jura Museum in Eichstätt, Germany, it is the smallest known specimen and has the second best head. It is possibly a separate genus (Jurapteryx recurva) or species (A. recurva). The Solnhofen Specimen (unnumbered specimen) was discovered in the 1970s near Eichstätt, Germany, and described in 1988 by Wellnhofer. Currently located at the Bürgermeister Müller Museum in Solnhofen, it originally was classified as Compsognathus by an amateur collector, the same mayor Friedrich Müller after which the museum is named. It is the largest specimen known and may belong to a separate genus and species, Wellnhoferia grandis. It is missing only portions of the neck, tail, backbone, and head. The Munich Specimen (BSP 1999 I 50, formerly known as the Solenhofer Aktien Verein Specimen) was discovered on 3 August 1992 near Langenaltheim and described in 1993 by Wellnhofer. It is currently located at the Paläontologisches Museum München in Munich, to which it was sold in 1999 for 1.9 million Deutschmark. What was initially believed to be a bony sternum turned out to be part of the coracoid, but more recent studies suggest it belongs to A. siemensii. After a lengthy period in a closed private collection, it was moved to the Museum of Evolution at Knuthenborg Safaripark (Denmark) in 2022, where it has since been on display and also been made available for researchers. Another fragmentary fossil was found in 2000. It is in private possession and, since 2004, on loan to the Bürgermeister Müller Museum in Solnhofen, so it is called the Bürgermeister Müller Specimen; the institute itself officially refers to it as the "Exemplar of the families Ottman & Steil, Solnhofen". As the fragment represents the remains of a single wing of Archaeopteryx, it is colloquially known as "chicken wing". Long in a private collection in Switzerland, the Thermopolis Specimen (WDC CSG 100) was discovered in Bavaria and described in 2005 by Mayr, Pohl, and Peters. Donated to the Wyoming Dinosaur Center in Thermopolis, Wyoming, it has the best preserved head and feet; most of the neck and the lower jaw have not been preserved. The "Thermopolis" specimen was described on 2 December 2005 Science journal article as "A well preserved Archaeopteryx specimen with theropod features"; it shows that Archaeopteryx lacked a reversed toe—a universal feature of birds—limiting its ability to perch on branches and implying a terrestrial or trunk climbing lifestyle. Palaeontologists of the Ludwig Maximilian University of Munich studied the specimen, which revealed previously unknown features of the plumage, such as feathers on both the upper and lower legs and metatarsus, and the only preserved tail tip. A twelfth specimen had been discovered by an amateur collector in 2010 at the Schamhaupten quarry, but the finding was only announced in February 2014. It was scientifically described in 2018. It represents a complete and mostly articulated skeleton with skull. It is the only specimen lacking preserved feathers. It is from the Painten Formation and somewhat older than the other specimens. The existence of a thirteenth specimen (the Chicago specimen) was announced in 2024 by the Field Museum in Chicago, US. One of two specimens in an institution outside Europe, the specimen was originally identified in a private collection in Switzerland, and had been acquired by these collectors in 1990, prior to Germany's 2015 ban on exporting Archaeopteryx specimens. The specimen was acquired by the Field Museum in 2022, and went on public display in 2024 following two years of preparation. The specimen is to be studied by famed paleornithologist Jingmai O'Connor.
Двенадцатый образец был обнаружен любителем-коллекционером в 2010 году на каменоломне Шамхауптен, но об этом было объявлено только в феврале 2014 года. Он был научно описан в 2018 году. Он представляет собой полный и в основном сочлененный скелет с черепом. Это единственный образец, у которого отсутствуют сохранившиеся перья. Он происходит из формации Пайнтен и немного старше других образцов.
the ancient animal
Archaeopteryx (,/ɑːr//k//iː/'/ɒ//p//t//ər//ᵻ//k//s/; ), sometimes referred to by its German name, "Urvogel" ( Primeval Bird) is a genus of bird like dinosaurs. The name derives from the ancient Greek (archaīos), meaning "ancient", and (ptéryx), meaning "feather" or "wing". Between the late 19th century and the early 21st century, Archaeopteryx was generally accepted by palaeontologists and popular reference books as the oldest known bird (member of the group Avialae). Older potential avialans have since been identified, including Anchiornis, Xiaotingia, and Aurornis. Archaeopteryx lived in the Late Jurassic around 150 million years ago, in what is now southern Germany, during a time when Europe was an archipelago of islands in a shallow warm tropical sea, much closer to the equator than it is now. Similar in size to a Eurasian magpie, with the largest individuals possibly attaining the size of a raven, Most of these twelve fossils include impressions of feathers. Because these feathers are of an advanced form (flight feathers), these fossils are evidence that the evolution of feathers began before the Late Jurassic. It is currently located at the Natural History Museum of Berlin. Though it was the initial holotype, there were indications that it might not have been from the same animal as the body fossils. This conclusion was challenged in 2020 as being unlikely; the feather was identified on the basis of morphology as most likely having been an upper major primary covert feather.) to the Natural History Museum in London, where it remains. The Haarlem Specimen (TM 6428/29, also known as the Teylers Specimen) was discovered in 1855 near italics=unset, Germany, and described as a Pterodactylus crassipes in 1857 by Meyer. It was reclassified in 1970 by John Ostrom and is currently located at the Teylers Museum in Haarlem, the Netherlands. It was the very first specimen found, but was incorrectly classified at the time. It is also one of the least complete specimens, consisting mostly of limb bones, isolated cervical vertebrae, and ribs. In 2017 it was named as a separate genus Ostromia, considered more closely related to Anchiornis from China. The Eichstätt Specimen (JM 2257) was discovered in 1951 near Workerszell, Germany, and described by Peter Wellnhofer in 1974. Currently located at the Jura Museum in Eichstätt, Germany, it is the smallest known specimen and has the second best head. It is possibly a separate genus (Jurapteryx recurva) or species (A. recurva). The Solnhofen Specimen (unnumbered specimen) was discovered in the 1970s near Eichstätt, Germany, and described in 1988 by Wellnhofer. Currently located at the Bürgermeister Müller Museum in Solnhofen, it originally was classified as Compsognathus by an amateur collector, the same mayor Friedrich Müller after which the museum is named. It is the largest specimen known and may belong to a separate genus and species, Wellnhoferia grandis. It is missing only portions of the neck, tail, backbone, and head. The Munich Specimen (BSP 1999 I 50, formerly known as the Solenhofer Aktien Verein Specimen) was discovered on 3 August 1992 near Langenaltheim and described in 1993 by Wellnhofer. It is currently located at the Paläontologisches Museum München in Munich, to which it was sold in 1999 for 1.9 million Deutschmark. What was initially believed to be a bony sternum turned out to be part of the coracoid, but more recent studies suggest it belongs to A. siemensii. After a lengthy period in a closed private collection, it was moved to the Museum of Evolution at Knuthenborg Safaripark (Denmark) in 2022, where it has since been on display and also been made available for researchers. Another fragmentary fossil was found in 2000. It is in private possession and, since 2004, on loan to the Bürgermeister Müller Museum in Solnhofen, so it is called the Bürgermeister Müller Specimen; the institute itself officially refers to it as the "Exemplar of the families Ottman & Steil, Solnhofen". As the fragment represents the remains of a single wing of Archaeopteryx, it is colloquially known as "chicken wing". Long in a private collection in Switzerland, the Thermopolis Specimen (WDC CSG 100) was discovered in Bavaria and described in 2005 by Mayr, Pohl, and Peters. Donated to the Wyoming Dinosaur Center in Thermopolis, Wyoming, it has the best preserved head and feet; most of the neck and the lower jaw have not been preserved. The "Thermopolis" specimen was described on 2 December 2005 Science journal article as "A well preserved Archaeopteryx specimen with theropod features"; it shows that Archaeopteryx lacked a reversed toe—a universal feature of birds—limiting its ability to perch on branches and implying a terrestrial or trunk climbing lifestyle. Palaeontologists of the Ludwig Maximilian University of Munich studied the specimen, which revealed previously unknown features of the plumage, such as feathers on both the upper and lower legs and metatarsus, and the only preserved tail tip. A twelfth specimen had been discovered by an amateur collector in 2010 at the Schamhaupten quarry, but the finding was only announced in February 2014. It was scientifically described in 2018. It represents a complete and mostly articulated skeleton with skull. It is the only specimen lacking preserved feathers. It is from the Painten Formation and somewhat older than the other specimens. The existence of a thirteenth specimen (the Chicago specimen) was announced in 2024 by the Field Museum in Chicago, US. One of two specimens in an institution outside Europe, the specimen was originally identified in a private collection in Switzerland, and had been acquired by these collectors in 1990, prior to Germany's 2015 ban on exporting Archaeopteryx specimens. The specimen was acquired by the Field Museum in 2022, and went on public display in 2024 following two years of preparation. The specimen is to be studied by famed paleornithologist Jingmai O'Connor.
Существование тринадцатого образца (Чикагский образец) было объявлено в 2024 году Музеем поля в Чикаго, США. Один из двух образцов в учреждении за пределами Европы, образец был первоначально идентифицирован в частной коллекции в Швейцарии и был приобретен этими коллекционерами в 1990 году, до запрета Германии в 2015 году на экспорт образцов археоптерикса. Образец был приобретен Музеем поля в 2022 году и был выставлен на всеобщее обозрение в 2024 году после двух лет подготовки. Образец будет изучен известным палеорнитологом Цзинмай О'Коннор.
the ancient animal
Archaeopteryx (,/ɑːr//k//iː/'/ɒ//p//t//ər//ᵻ//k//s/; ), sometimes referred to by its German name, "Urvogel" ( Primeval Bird) is a genus of bird like dinosaurs. The name derives from the ancient Greek (archaīos), meaning "ancient", and (ptéryx), meaning "feather" or "wing". Between the late 19th century and the early 21st century, Archaeopteryx was generally accepted by palaeontologists and popular reference books as the oldest known bird (member of the group Avialae). Older potential avialans have since been identified, including Anchiornis, Xiaotingia, and Aurornis. Archaeopteryx lived in the Late Jurassic around 150 million years ago, in what is now southern Germany, during a time when Europe was an archipelago of islands in a shallow warm tropical sea, much closer to the equator than it is now. Similar in size to a Eurasian magpie, with the largest individuals possibly attaining the size of a raven, Most of these twelve fossils include impressions of feathers. Because these feathers are of an advanced form (flight feathers), these fossils are evidence that the evolution of feathers began before the Late Jurassic. It is currently located at the Natural History Museum of Berlin. Though it was the initial holotype, there were indications that it might not have been from the same animal as the body fossils. This conclusion was challenged in 2020 as being unlikely; the feather was identified on the basis of morphology as most likely having been an upper major primary covert feather.) to the Natural History Museum in London, where it remains. The Haarlem Specimen (TM 6428/29, also known as the Teylers Specimen) was discovered in 1855 near italics=unset, Germany, and described as a Pterodactylus crassipes in 1857 by Meyer. It was reclassified in 1970 by John Ostrom and is currently located at the Teylers Museum in Haarlem, the Netherlands. It was the very first specimen found, but was incorrectly classified at the time. It is also one of the least complete specimens, consisting mostly of limb bones, isolated cervical vertebrae, and ribs. In 2017 it was named as a separate genus Ostromia, considered more closely related to Anchiornis from China. The Eichstätt Specimen (JM 2257) was discovered in 1951 near Workerszell, Germany, and described by Peter Wellnhofer in 1974. Currently located at the Jura Museum in Eichstätt, Germany, it is the smallest known specimen and has the second best head. It is possibly a separate genus (Jurapteryx recurva) or species (A. recurva). The Solnhofen Specimen (unnumbered specimen) was discovered in the 1970s near Eichstätt, Germany, and described in 1988 by Wellnhofer. Currently located at the Bürgermeister Müller Museum in Solnhofen, it originally was classified as Compsognathus by an amateur collector, the same mayor Friedrich Müller after which the museum is named. It is the largest specimen known and may belong to a separate genus and species, Wellnhoferia grandis. It is missing only portions of the neck, tail, backbone, and head. The Munich Specimen (BSP 1999 I 50, formerly known as the Solenhofer Aktien Verein Specimen) was discovered on 3 August 1992 near Langenaltheim and described in 1993 by Wellnhofer. It is currently located at the Paläontologisches Museum München in Munich, to which it was sold in 1999 for 1.9 million Deutschmark. What was initially believed to be a bony sternum turned out to be part of the coracoid, but more recent studies suggest it belongs to A. siemensii. After a lengthy period in a closed private collection, it was moved to the Museum of Evolution at Knuthenborg Safaripark (Denmark) in 2022, where it has since been on display and also been made available for researchers. Another fragmentary fossil was found in 2000. It is in private possession and, since 2004, on loan to the Bürgermeister Müller Museum in Solnhofen, so it is called the Bürgermeister Müller Specimen; the institute itself officially refers to it as the "Exemplar of the families Ottman & Steil, Solnhofen". As the fragment represents the remains of a single wing of Archaeopteryx, it is colloquially known as "chicken wing". Long in a private collection in Switzerland, the Thermopolis Specimen (WDC CSG 100) was discovered in Bavaria and described in 2005 by Mayr, Pohl, and Peters. Donated to the Wyoming Dinosaur Center in Thermopolis, Wyoming, it has the best preserved head and feet; most of the neck and the lower jaw have not been preserved. The "Thermopolis" specimen was described on 2 December 2005 Science journal article as "A well preserved Archaeopteryx specimen with theropod features"; it shows that Archaeopteryx lacked a reversed toe—a universal feature of birds—limiting its ability to perch on branches and implying a terrestrial or trunk climbing lifestyle. Palaeontologists of the Ludwig Maximilian University of Munich studied the specimen, which revealed previously unknown features of the plumage, such as feathers on both the upper and lower legs and metatarsus, and the only preserved tail tip. A twelfth specimen had been discovered by an amateur collector in 2010 at the Schamhaupten quarry, but the finding was only announced in February 2014. It was scientifically described in 2018. It represents a complete and mostly articulated skeleton with skull. It is the only specimen lacking preserved feathers. It is from the Painten Formation and somewhat older than the other specimens. The existence of a thirteenth specimen (the Chicago specimen) was announced in 2024 by the Field Museum in Chicago, US. One of two specimens in an institution outside Europe, the specimen was originally identified in a private collection in Switzerland, and had been acquired by these collectors in 1990, prior to Germany's 2015 ban on exporting Archaeopteryx specimens. The specimen was acquired by the Field Museum in 2022, and went on public display in 2024 following two years of preparation. The specimen is to be studied by famed paleornithologist Jingmai O'Connor.
Подлинность
Начиная с 1985 года, группа энтузиастов, включавшая астронома Фреда Хойла и физика Ли Спетнера, опубликовала серию статей, в которых утверждалось, что перья на берлинских и лондонских экземплярах археоптерикса были подделаны. Археоптерикс был размером примерно с ворона, и хотя перья археоптерикса задокументированы меньше, чем другие его особенности, по своей структуре они были очень похожи на перья современных птиц.
Оперение
У образцов археоптерикса наиболее заметны их хорошо развитые маховые перья. Они были заметно асимметричны и демонстрировали структуру маховых перьев у современных птиц, причем опахала были укреплены системой бородко-бородчатых зацепок. Это означает, что кожа уже была смягчена и стала рыхлой, что подтверждается тем фактом, что у некоторых экземпляров маховые перья начинали отсоединяться в месте внедрения в осадочную породу. Таким образом, предполагается, что соответствующие экземпляры какое-то время перед погребением перемещались по морскому дну в неглубокой воде, при этом перья головы и верхней части шеи опадали, а более прочно прикрепленные хвостовые перья оставались на месте. В 2011 году аспирант Райан Карни и его коллеги провели первое исследование окраски оперения экземпляра археоптерикса. В исследовании 2013 года, опубликованном в журнале "Journal of Analytical Atomic Spectrometry", новый анализ перьев археоптерикса показал, что у животного могло быть сложное оперение, сочетающее светлые и темные цвета, с более интенсивной пигментацией на дистальных концах и внешних опахалах. Этот анализ распределения цвета основывался главным образом на распределении сульфатов в ископаемом. Автор предыдущего исследования окраски археоптерикса выступил против интерпретации таких биомаркеров как индикатора эумеланина в полном экземпляре археоптерикса. Карни и другие коллеги также оспорили интерпретацию сульфатов и микроэлементов в исследовании 2013 года, а в исследовании 2020 года, опубликованном в Scientific Reports, было показано, что изолированное кроющее перо было полностью матово-черным (в отличие от черно-белого или переливающегося) и что остальные "паттерны оперения археоптерикса остаются неизвестными". Это предложение было поддержано ICZN после четырех лет дебатов, и лондонский образец был назначен неотипом 3 октября 2011 года. Ниже представлена кладограмма, опубликованная в 2013 году Годфруа и др. Филогенетические исследования, проведенные Сентером и др. (2012) и Тернером, Маковицки и Нореллом (2012) также показали, что археоптерикс более тесно связан с современными птицами, чем с дромеозавридами и троодонтидами. С другой стороны, Годфруа и др. (2013) обнаружили, что археоптерикс более тесно связан с дромеозавридами и троодонтидами в анализе, включенном в их описание Eosinopteryx brevipenna. Авторы использовали модифицированную версию матрицы из исследования, описывающего Xiaotingia, добавив Jinfengopteryx elegans и Eosinopteryx brevipenna, а также четыре дополнительных признака, связанных с развитием оперения. В отличие от анализа из описания Xiaotingia, анализ, проведенный Годфруа и др., не показал, что археоптерикс особенно тесно связан с Anchiornis и Xiaotingia, которые были восстановлены как базальные троодонтиды. Агнолин и Новас (2013) обнаружили, что археоптерикс и (возможно, синонимичный) Wellnhoferia образуют кладу-сестру к линии, включающей Jeholornis и Pygostylia, при этом Microraptoria, Unenlagiinae и клада, содержащая Anchiornis и Xiaotingia, последовательно являются более отдаленными группами по отношению к Avialae (определенной авторами как клада, происходящая от последнего общего предка Archaeopteryx и Aves). Другое филогенетическое исследование Годфруа и др., использующее более инклюзивную матрицу, чем та, которая была использована в анализе в описании Eosinopteryx brevipenna, также показало, что археоптерикс является членом Avialae (определенной авторами как наиболее инклюзивная клада, содержащая Passer domesticus, но не Dromaeosaurus albertensis или Troodon formosus). Было обнаружено, что археоптерикс образует класс у основания Avialae вместе с Xiaotingia, Anchiornis и Aurornis. По сравнению с археоптериксом, Xiaotingia оказался более тесно связан с существующими птицами, в то время как Anchiornis и Aurornis были более отдаленными. Ван и др. (2018) и Хартман и др. (2019) обнаружили, что археоптерикс был динонихозавром, а не птицеловом. В частности, он и близкородственные таксоны считались базальными динонихозаврами, при этом дромеозавриды и троодонтиды вместе образовывали параллельную линию внутри группы. Поскольку Хартман и др. обнаружили, что археоптерикс изолирован в группе нелетающих динонихозавров (иначе говоря, считаемых "анхиорнитидами"), они считают весьма вероятным, что это животное развило полет независимо от предков птиц (и от Microraptor и Yi). Следующая кладограмма иллюстрирует их гипотезу относительно положения археоптерикса: авторы, однако, обнаружили, что принадлежность археоптерикса к Avialae была лишь немного менее вероятной, чем эта гипотеза, и столь же вероятной, как сестринские отношения между Archaeopterygidae и Troodontidae. Крылья археоптерикса были относительно большими, что привело бы к низкой скорости сваливания и уменьшению радиуса поворота. Короткая и округлая форма крыльев увеличила бы сопротивление, но также могла улучшить его способность летать в загроможденной среде, такой как деревья и кустарники (подобная форма крыльев встречается у птиц, летающих сквозь деревья и кустарники, таких как вороны и фазаны). Наличие "задних крыльев" – асимметричных маховых перьев, растущих из ног, подобных тем, что встречаются у дромеозавридов, таких как Microraptor, также способствовало бы маневренности археоптерикса в воздухе. Первое подробное исследование задних крыльев, проведенное Лонгричем в 2006 году, показало, что они составляли до 12% от общей площади крыла. Это могло снизить скорость сваливания до 6% и радиус поворота до 12%. Степень асимметрии, наблюдаемая у археоптерикса, более типична для медленно летающих птиц, чем для нелетающих. В 2010 году Роберт Л. Наддс и Гарет Дж. Дайк в журнале Science опубликовали статью, в которой они проанализировали стержни первичных перьев Confuciusornis и археоптерикса. Анализ показал, что стержни у этих двух родов были тоньше и слабее, чем у современных птиц, относительно массы тела. Авторы пришли к выводу, что археоптерикс и Confuciusornis не могли использовать взмахивающим полетом. Это исследование подверглось критике со стороны Филипа Дж. Карри и Луиса Чиаппе. Чиаппе предположил, что трудно измерить стержни окаменелых перьев, а Карри предположил, что археоптерикс и Confuciusornis должны были уметь летать в какой-то степени, поскольку их окаменелости сохранились в том, что считается морскими или озерными отложениями, что указывает на то, что они должны были уметь летать над глубокой водой. Грегори Пол также не согласился с исследованием, заявив в своем ответе в 2010 году, что Наддс и Дайк переоценили массу этих ранних птиц, и что более точные оценки массы позволяют осуществлять полет даже с относительно узкими стержнями. Наддс и Дайк предположили массу в для мюнхенского экземпляра археоптерикса, молодого детеныша, основываясь на опубликованных оценках массы более крупных экземпляров. Пол утверждал, что более разумная оценка массы тела для мюнхенского экземпляра составляет около Пол также раскритиковал измерения самих стержней, отметив, что перья в мюнхенском экземпляре плохо сохранились. Наддс и Дайк сообщили о диаметре для самого длинного махового пера, что Пол не смог подтвердить, используя фотографии. Пол измерил некоторые из внутренних маховых перьев, обнаружив стержни в поперечнике. Несмотря на эту критику, Наддс и Дайк остались при своих первоначальных выводах. Они заявили, что утверждение Пола о том, что взрослый археоптерикс был бы лучшим летуном, чем юный мюнхенский экземпляр, было сомнительным. По их мнению, это потребовало бы еще более толстого стержня, доказательств которого пока не представлено. Другая возможность заключается в том, что они не достигли истинного полета, а вместо этого использовали свои крылья в качестве вспомогательного средства для дополнительной подъемной силы при беге по воде, как это делает василиск, что может объяснить их присутствие в озерных и морских отложениях (см. Происхождение птичьего полета). В 2004 году ученые, анализирующие подробное КТ-сканирование мозгового черепа лондонского археоптерикса, пришли к выводу, что его мозг был значительно больше, чем у большинства динозавров, что указывает на то, что он обладал развитым...
The authors, however, found that the Archaeopteryx being an avialan was only slightly less likely than this hypothesis, and as likely as Archaeopterygidae and Troodontidae being sister clades. Archaeopteryx wings were relatively large, which would have resulted in a low stall speed and reduced turning radius. The short and rounded shape of the wings would have increased drag, but also could have improved its ability to fly through cluttered environments such as trees and brush (similar wing shapes are seen in birds that fly through trees and brush, such as crows and pheasants). The presence of "hind wings", asymmetrical flight feathers stemming from the legs similar to those seen in dromaeosaurids such as Microraptor, also would have added to the aerial mobility of Archaeopteryx. The first detailed study of the hind wings by Longrich in 2006, suggested that the structures formed up to 12% of the total airfoil. This would have reduced stall speed by up to 6% and turning radius by up to 12%. The degree of asymmetry seen in Archaeopteryx is more typical for slow flyers than for flightless birds. In 2010, Robert L. Nudds and Gareth J. Dyke in the journal Science published a paper in which they analysed the rachises of the primary feathers of Confuciusornis and Archaeopteryx. The analysis suggested that the rachises on these two genera were thinner and weaker than those of modern birds relative to body mass. The authors determined that Archaeopteryx and Confuciusornis, were unable to use flapping flight. This study was criticized by Philip J. Currie and Luis Chiappe. Chiappe suggested that it is difficult to measure the rachises of fossilized feathers, and Currie speculated that Archaeopteryx and Confuciusornis must have been able to fly to some degree, as their fossils are preserved in what is believed to have been marine or lake sediments, suggesting that they must have been able to fly over deep water. Gregory Paul also disagreed with the study, arguing in a 2010 response that Nudds and Dyke had overestimated the masses of these early birds, and that more accurate mass estimates allowed powered flight even with relatively narrow rachises. Nudds and Dyke had assumed a mass of for the Munich specimen Archaeopteryx, a young juvenile, based on published mass estimates of larger specimens. Paul argued that a more reasonable body mass estimate for the Munich specimen is about Paul also criticized the measurements of the rachises themselves, noting that the feathers in the Munich specimen are poorly preserved. Nudds and Dyke reported a diameter of for the longest primary feather, which Paul could not confirm using photographs. Paul measured some of the inner primary feathers, finding rachises across. Despite these criticisms, Nudds and Dyke stood by their original conclusions. They claimed that Paul's statement, that an adult Archaeopteryx would have been a better flyer than the juvenile Munich specimen, was dubious. This, they reasoned, would require an even thicker rachis, evidence for which has not yet been presented. Another possibility is that they had not achieved true flight, but instead used their wings as aids for extra lift while running over water after the fashion of the basilisk lizard, which could explain their presence in lake and marine deposits (see Origin of avian flight). In 2004, scientists analysing a detailed CT scan of the braincase of the London Archaeopteryx concluded that its brain was significantly larger than that of most dinosaurs, indicating that it possessed the brain size necessary for flying. The overall brain anatomy was reconstructed using the scan. The reconstruction showed that the regions associated with vision took up nearly one third of the brain. Other well developed areas involved hearing and muscle coordination. Newer research shows that while the Archaeopteryx brain was more complex than that of more primitive theropods, it had a more generalized brain volume among Maniraptora dinosaurs, even smaller than that of other non avian dinosaurs in several instances, which indicates the neurological development required for flight was already a common trait in the maniraptoran clade. Recent studies of flight feather barb geometry reveal that modern birds possess a larger barb angle in the trailing vane of the feather, whereas Archaeopteryx lacks this large barb angle, indicating potentially weak flight abilities. Archaeopteryx continues to play an important part in scientific debates about the origin and evolution of birds. Some scientists see it as a semi arboreal climbing animal, following the idea that birds evolved from tree dwelling gliders (the "trees down" hypothesis for the evolution of flight proposed by O. C. Marsh). Other scientists see Archaeopteryx as running quickly along the ground, supporting the idea that birds evolved flight by running (the "ground up" hypothesis proposed by Samuel Wendell Williston). Still others suggest that Archaeopteryx might have been at home both in the trees and on the ground, like modern crows, and this latter view is what currently is considered best supported by morphological characters. Altogether, it appears that the species was not particularly specialized for running on the ground or for perching. A scenario outlined by Elżanowski in 2002 suggested that Archaeopteryx used its wings mainly to escape predators by glides punctuated with shallow downstrokes to reach successively higher perches, and alternatively, to cover longer distances (mainly) by gliding down from cliffs or treetops. This study on Archaeopteryxs bone histology identified biomechanical and physiological adaptations exhibited by modern volant birds that perform intermittent flapping, such as pheasants and other burst flyers. Studies of Archaeopteryx's feather sheaths revealed that like modern birds, it had a center out, flight related molting strategy. As it was a weak flier, this was extremely advantageous in preserving its maximum flight performance.
Рост
Гистологическое исследование, проведенное Эриксоном, Нореллом, Чжунгу и другими в 2009 году, показало, что археоптерикс рос относительно медленно по сравнению с современными птицами, вероятно, из-за того, что внешние слои костей археоптерикса выглядят слабо васкуляризованными. Одной из немногих современных птиц, демонстрирующих замедленный рост, является нелетающий киви, и авторы предположили, что у археоптерикса и киви был схожий уровень базального метаболизма.
Палеоэкология
Богатство и разнообразие известняков Солнхофена, в которых найдены все известные экземпляры археоптерикса, пролили свет на древнюю юрскую Баварию, разительно отличающуюся от современной. Широта была схожа с флоридской, однако климат, вероятно, был более сухим, что подтверждается окаменелостями растений, адаптированных к засушливым условиям, и отсутствием наземных отложений, характерных для речных систем. Хотя свидетельства о растительности немногочисленны, среди них встречаются цикадовые и хвойные растения, а также большое количество насекомых, мелкие ящерицы, птерозавры и компсогнаты. Отсутствие деревьев не исключает древесный образ жизни археоптерикса, поскольку некоторые виды птиц живут исключительно в низкорослых кустарниках. Различные аспекты морфологии археоптерикса указывают на возможность как древесного, так и наземного образа жизни, включая длину ног и удлинение стоп; некоторые исследователи полагают, что он, вероятно, был всеядным, способным питаться как в кустарниках, так и на открытой местности, а также вдоль берегов лагуны.