Генотип пен ортаның өзара әрекеттесуінен туындайтын фенотиптік өрнектер

Pattern of phenotypic expression caused by a given genotype across a range of environments
In ecology and genetics, a reaction norm, also called a norm of reaction, describes the pattern of phenotypic expression of a single genotype across a range of environments. One use of reaction norms is in describing how different species—especially related species—respond to varying environments. But differing genotypes within a single species may also show differing reaction norms relative to a particular phenotypic trait and environment variable. For every genotype, phenotypic trait, and environmental variable, a different reaction norm can exist; in other words, an enormous complexity can exist in the interrelationships between genetic and environmental factors in determining traits. The concept was introduced by Richard Woltereck in 1909.

Scientifically analyzing norms of reaction in natural populations can be very difficult, simply because natural populations of sexually reproductive organisms usually do not have cleanly separated or superficially identifiable genetic distinctions. However, seed crops produced by humans are often engineered to contain specific genes, and in some cases seed stocks consist of clones. Accordingly, distinct seed lines present ideal examples of differentiated norms of reaction. In fact, agricultural companies market seeds for use in particular environments based on exactly this. Suppose the seed line A contains an allele a, and a seed line B of the same crop species contains an allele b, for the same gene. With these controlled genetic groups, we might cultivate each variety (genotype) in a range of environments. This range might be either natural or controlled variations in environment. For example, an individual plant might receive either more or less water during its growth cycle, or the average temperature the plants are exposed to might vary across a range. A simplification of the norm of reaction might state that seed line A is good for "high water conditions" while a seed line B is good for "low water conditions". But the full complexity of the norm of reaction is a function, for each genotype, relating environmental factor to phenotypic trait. By controlling for or measuring actual environments across which monoclonal seeds are cultivated, one can concretely observe norms of reaction. Normal distributions, for example, are common. Of course, the distributions need not be bell curves.

One advantage of plants is that the same genotype, such as a recombinant inbred line (RIL), can be repeatedly evaluated in multiple environments, or a multi environmental trial (MET). The reaction norm can then be explored based on the geographic location, mean trait value summarized from the whole population at each environment, or an explicit performance free index capturing relevant environment inputs.