Семейство куликов Scolopacidae: разнообразие и особенности

Family of birds
the bird

Scolopacidae is a large family of shorebirds, or waders, which mainly includes many species known as sandpipers, but also others such as woodcocks, curlews and snipes. The majority of these species eat small invertebrates picked out of the mud or soil. Different lengths of bills enable different species to feed in the same habitat, particularly on the coast, without direct competition for food. Sandpipers have long bodies and legs, and narrow wings. Most species have a narrow bill, but otherwise the form and length are quite variable. They are small to medium sized birds, measuring in length. The bills are sensitive, allowing the birds to feel the mud and sand as they probe for food. They generally have dull plumage, with cryptic brown, grey, or streaked patterns, although some display brighter colours during the breeding season. Most species nest in open areas, and defend their territories with aerial displays. The nest itself is a simple scrape in the ground, in which the bird typically lays three or four eggs. The young of most species are precocial. The family contains 98 extant or recently extinct species divided into 15 genera. For more details, see the article List of sandpiper species. The following genus level cladogram of the Scolopacidae is based on a study by David Černý and Rossy Natale that was published in 2022.

The sandpipers exhibit considerable range in size and appearance, the wide range of body forms reflecting a wide range of ecological niches. Sandpipers range in size from the least sandpiper, at as little as and in length, to the Far Eastern curlew, at up to in length, and the Eurasian curlew, at up to Within species there is considerable variation in patterns of sexual dimorphism. Males are larger than females in ruffs and several sandpipers, but are smaller than females in the knots, curlews, phalaropes and godwits. The sexes are similarly sized in the snipes, woodcock and tringine sandpipers. Compared to the other large family of wading birds, the plovers (Charadriidae), they tend to have smaller eyes, more slender heads, and longer thinner bills. Some are quite long legged, and most species have three forward pointing toes with a smaller hind toe (the exception is the sanderling, which lacks a hind toe). Like all birds, the bills of sandpipers are capable of cranial kinesis, literally being able to move the bones of the skull (other than the obvious movement of the lower jaw) and specifically bending the upper jaw without opening the entire jaw, an act known as rhynchokinesis. It has been hypothesized this helps when probing by allowing the bill to be partly opened with less force and improving manipulation of prey items in the substrate. Rhynchokinesis is also used by sandpipers feeding on prey in water to catch and manipulate prey.

The sandpipers have a cosmopolitan distribution, occurring across most of the world's land surfaces except for Antarctica and the driest deserts. A majority of the family breed at moderate to high latitudes in the Northern Hemisphere, in fact accounting for the most northerly breeding birds in the world. Only a few species breed in tropical regions, ten of which are snipes and woodcocks and the remaining species being the unusual Tuamotu sandpiper, which breeds in French Polynesia (although prior to the arrival of humans in the Pacific there were several other closely related species of Polynesian sandpiper). In species where only a single parent incubates the eggs, during the night the parent sits on the eggs nearly continuously and then during the warmest part of a day leaves the nest for short feeding bouts. Chicks hatch after about three weeks of incubation and are able to walk and forage within a few hours of hatching. A single parent or both parents guide and brood the chicks.