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The iOS version of Ascendancy was a Universal app, meaning it was designed for both the iPad and the iPhone / iPod touch platforms. The original Ascendancy was released during a golden age of 4X space games in the mid 1990s. The player does this by building structures and space ships, researching technology and growing stronger through war or diplomacy. Players begin with a single colony planet which must be developed. They can select several types of buildings to construct on the planet's surface and can eventually construct spaceships and leave the planet altogether. Ships travel to other solar systems along determined paths, called star lanes. Spaceships can peacefully colonize unoccupied planets or invade inhabited planets and usurp structures. They can also do battle with enemy planets or other ships. Ascendancy's primary display is a three dimensional rotatable model of the star cluster, which details the star cluster's solar systems, starlanes, and civilizations. The player can "zoom in" to view rotatable models of solar systems (including planets and ships); view a planet's surface and use various screens to manage their fleet, planets and research. This play style is similar to the Master of Orion series.

On each planet the player must manage three areas of development: Research, Industry and Prosperity. Research points are shared among all colonies and allow the player to learn new technology, such as advanced building structures or spaceship components. Industry, meanwhile, is local to each colony and determines how quickly that planet can complete projects. Prosperity determines how quickly a planet's population will grow, providing additional workers for future construction. Players begin with the ability to construct buildings for each of these areas and can discover more as the game progresses. Planet topography is expressed as various colored squares. White squares are general purpose, where any structure can be built. Green squares increase the output of Prosperity producing buildings, while red squares improve Industry and blue squares improve Research. There are also black squares, which are initially uninhabitable but can be used later in the game as the player researches technology. Planets have ten orbital squares that can be used to build shipyards, planetary defenses, space ships, and other space structures. As the player explores other planets they may occasionally find ruins from an ancient civilization. These ruins can be harvested to gain undiscovered technologies, which can significantly alter the game.

Ascendancy contains a Diplomacy screen where players can offer alliances to other species, declare war and exchange information on star lanes, planets or research. Players can request military assistance from their allies or attempt to end declarations of war with hostile factions. There is also an Intelligence display where the player can view information on the relative power of known species.

When starting a new game players select several parameters, including their species, a star density, a political atmosphere and the number of opponents. There are 21 species to choose from in Ascendancy, each of which has a unique special ability. Some abilities can be used at regular intervals, such as the Capelon's ability to make their colonies temporarily invulnerable. Other abilities, such as the Chronomyst traveling quickly through star lanes, are continually active. All opponent species (numbering from two to six) are controlled by a computer AI and are selected randomly. The star density in each game determines how many solar systems exist in the star cluster the higher the density the more planets that are available to colonize. Star density affects both the length of the game, as large star clusters will take longer to conquer, as well as affecting how quickly species will encounter each other and how much room they have to grow. Once a density is selected the particular type of each star system is generated randomly at the start of the game. Selecting a political atmosphere of Peaceful, Neutral or Hostile determines the attitude of computer opponents when they have a colony in the same system as another species. Even though the player has control over several things, the combination of random opponent species, random planet topography and random star types leads to a large variety of games.

Several of the species in Ascendancy describe motivations for reaching to the stars. Some species are peaceful diplomats; some are curious explorers; and others want to kill or conquer other forms of life. However, plot in Ascendancy is thin. Players are free to imagine a detailed storyline but most plot is restricted to a single introductory screen for each species and a synopsis given at the end of each game.

The Logic Factory said that inspiration for Ascendancy came from "[having] read the fantastic books or watched some of the great shows that have explored the possibility of human contact with intelligent alien life. We do believe that there is a high likelihood that among the many billions of stars and planets just in our own Milky Way, other life is out there. This [game] is our own way of being part of that inevitable adventure."

In 2010 the Logic Factory updated their website and announced that a remake of Ascendancy would be released for the iPhone and iPad, intended as "a fully upgraded remake tailored for modern wireless gaming." An iOS version of Ascendancy was released on the Apple app store on Jan 05, 2011. It has since been removed.

Some reviewers sang high praises for Ascendancy upon release the game won the Codie award for Best Strategy Software in 1996, while Next Generation Magazine called Ascendancy "one of the best space strategy sims" and said "the variety of species will give you hours of replay value." In his 1995 PC Gamer review William Trotter called Ascendancy "simply the best game of its type yet published" and gave it the PC Gamer Editor's Choice award. Bruce Geryk said "the graphics were excellent, and the method of ship design was quite engaging," while Andy Bucher of PC Gamer UK said "it looks great everything looks slick, clean and well designed Ascendancy mixes just about the right level of complexity and challenge with an intuitive interface, great graphics and the odd spark of humour." Bucher even noted that Ascendancy had "one of the best tutorials [he'd] ever seen", and said the game's "attention to detail and ease of use is much rarer than it should be." The iOS version of Ascendancy released in 2011 included the AI from this patch, as well as other modifications.