Жұмырқа қаңқасы және оның құрылысы

Bony structure that forms the head in vertebrates
the skulls of all animals including humans

The skull is a bone protective cavity for the brain. The skull is composed of four types of bone i. e., cranial bones, facial bones, ear ossicles and hyoid bone, however two parts are more prominent: the cranium (: craniums or crania) and the mandible. In humans, these two parts are the neurocranium (braincase) and the viscerocranium (facial skeleton) that includes the mandible as its largest bone. The skull forms the anterior most portion of the skeleton and is a product of cephalisation—housing the brain, and several sensory structures such as the eyes, ears, nose, and mouth. In humans, these sensory structures are part of the facial skeleton. Functions of the skull include protection of the brain, fixing the distance between the eyes to allow stereoscopic vision, and fixing the position of the ears to enable sound localisation of the direction and distance of sounds. In some animals, such as horned ungulates (mammals with hooves), the skull also has a defensive function by providing the mount (on the frontal bone) for the horns. The English word skull is probably derived from Old Norse skulle, while the Latin word cranium comes from the Greek root κρανίον (kranion). The human skull fully develops two years after birth. The junctions of the skull bones are joined by structures called sutures. The skull is made up of a number of fused flat bones, and contains many foramina, fossae, processes, and several cavities or sinuses. In zoology, there are openings in the skull called fenestrae.

The human skull is the bone structure that forms the head in the human skeleton. It supports the structures of the face and forms a cavity for the brain. Like the skulls of other vertebrates, it protects the brain from injury. The skull consists of three parts, of different embryological origin—the neurocranium, the sutures, and the facial skeleton. The neurocranium (or braincase) forms the protective cranial cavity that surrounds and houses the brain and brainstem. The upper areas of the cranial bones form the calvaria (skullcap). The membranous viscerocranium includes the mandible. The sutures are fairly rigid joints between bones of the neurocranium. The facial skeleton is formed by the bones supporting the face.

Except for the mandible, all of the bones of the skull are joined by sutures—synarthrodial (immovable) joints formed by bony ossification, with Sharpey's fibres permitting some flexibility. Sometimes there can be extra bone pieces within the suture known as Wormian bones or sutural bones. Most commonly these are found in the course of the lambdoid suture. The human skull is generally considered to consist of 22 bones—eight cranial bones and fourteen facial skeleton bones. In the neurocranium these are the occipital bone, two temporal bones, two parietal bones, the sphenoid, ethmoid and frontal bones. The bones of the facial skeleton (14) are the vomer, two inferior nasal conchae, two nasal bones, two maxilla, the mandible, two palatine bones, two zygomatic bones, and two lacrimal bones. Some sources count a paired bone as one, or the maxilla as having two bones (as its parts); some sources include the hyoid bone or the three ossicles of the middle ear, the malleus, incus, and stapes, but the overall general consensus of the number of bones in the human skull is the stated twenty two. Some of these bones—the occipital, parietal, frontal, in the neurocranium, and the nasal, lacrimal, and vomer, in the facial skeleton are flat bones.

The skull also contains sinuses, air filled cavities known as paranasal sinuses, and numerous foramina. The sinuses are lined with respiratory epithelium. Their known functions are the lessening of the weight of the skull, the aiding of resonance to the voice and the warming and moistening of the air drawn into the nasal cavity. The foramina are openings in the skull. The largest of these is the foramen magnum, of the occipital bone, that allows the passage of the spinal cord as well as nerves and blood vessels.

The many processes of the skull include the mastoid process and the zygomatic processes.

The jugal is a skull bone found in most reptiles, amphibians, and birds. In mammals, the jugal is often called the zygomatic bone or malar bone. The prefrontal bone is a bone separating the lacrimal and frontal bones in many tetrapod skulls.

The skull of fishes is formed from a series of only loosely connected bones. Lampreys and sharks only possess a cartilaginous endocranium, with both the upper and lower jaws being separate elements. Bony fishes have additional dermal bone, forming a more or less coherent skull roof in lungfish and holost fish. The lower jaw defines a chin. The simpler structure is found in jawless fish, in which the cranium is normally represented by a trough like basket of cartilaginous elements only partially enclosing the brain, and associated with the capsules for the inner ears and the single nostril. Distinctively, these fish have no jaws. In ray finned fish, there has also been considerable modification from the primitive pattern. The roof of the skull is generally well formed, and although the exact relationship of its bones to those of tetrapods is unclear, they are usually given similar names for convenience. Other elements of the skull, however, may be reduced; there is little cheek region behind the enlarged orbits, and little, if any bone in between them. The upper jaw is often formed largely from the premaxilla, with the maxilla itself located further back, and an additional bone, the symplectic, linking the jaw to the rest of the cranium. In living tetrapods, a great many of the original bones have either disappeared or fused into one another in various arrangements.

Birds have a diapsid skull, as in reptiles, with a prelacrimal fossa (present in some reptiles). The skull has a single occipital condyle. The skull consists of five major bones: the frontal (top of head), parietal (back of head), premaxillary and nasal (top beak), and the mandible (bottom beak). The skull of a normal bird usually weighs about 1% of the bird's total bodyweight. The eye occupies a considerable amount of the skull and is surrounded by a sclerotic eye ring, a ring of tiny bones. This characteristic is also seen in reptiles.

Living amphibians typically have greatly reduced skulls, with many of the bones either absent or wholly or partly replaced by cartilage. The geometry of the skull base and its fossae, the anterior, middle and posterior cranial fossae changes rapidly. The anterior cranial fossa changes especially during the first trimester of pregnancy and skull defects can often develop during this time. At birth, the human skull is made up of 44 separate bony elements. During development, many of these bony elements gradually fuse together into solid bone (for example, the frontal bone). The bones of the roof of the skull are initially separated by regions of dense connective tissue called fontanelles. There are six fontanelles: one anterior (or frontal), one posterior (or occipital), two sphenoid (or anterolateral), and two mastoid (or posterolateral). At birth, these regions are fibrous and moveable, necessary for birth and later growth. This growth can put a large amount of tension on the "obstetrical hinge", which is where the squamous and lateral parts of the occipital bone meet. A possible complication of this tension is rupture of the great cerebral vein. As growth and ossification progress, the connective tissue of the fontanelles is invaded and replaced by bone creating sutures. The five sutures are the two squamous sutures, one coronal, one lambdoid, and one sagittal suture. The posterior fontanelle usually closes by eight weeks, but the anterior fontanel can remain open up to eighteen months. The anterior fontanelle is located at the junction of the frontal and parietal bones; it is a "soft spot" on a baby's forehead. Careful observation will show that you can count a baby's heart rate by observing the pulse pulsing softly through the anterior fontanelle. The skull in the neonate is large in proportion to other parts of the body. The facial skeleton is one seventh of the size of the calvaria. (In the adult it is half the size). The base of the skull is short and narrow, though the inner ear is almost adult size.

Craniosynostosis is a condition in which one or more of the fibrous sutures in an infant skull prematurely fuses, and changes the growth pattern of the skull. Because the skull cannot expand perpendicular to the fused suture, it grows more in the parallel direction. A copper beaten skull is a phenomenon wherein intense intracranial pressure disfigures the internal surface of the skull. The name comes from the fact that the inner skull has the appearance of having been beaten with a ball peen hammer, such as is often used by coppersmiths. The condition is most common in children.

Injuries to the brain can be life threatening. Normally the skull protects the brain from damage through its high resistance to deformation; the skull is one of the least deformable structures found in nature, needing the force of about 1 ton to reduce its diameter by 1 cm. In some cases of head injury, however, there can be raised intracranial pressure through mechanisms such as a subdural haematoma. In these cases, the raised intracranial pressure can cause herniation of the brain out of the foramen magnum ("coning") because there is no space for the brain to expand; this can result in significant brain damage or death unless an urgent operation is performed to relieve the pressure. This is why patients with concussion must be watched extremely carefully. Repeated concussions can activate the structure of skull bones as the brain's protective covering. Dating back to Neolithic times, a skull operation called trepanning was sometimes performed. This involved drilling a burr hole in the cranium. Examination of skulls from this period reveals that the patients sometimes survived for many years afterward. It seems likely that trepanning was also performed purely for ritualistic or religious reasons. Nowadays this procedure is still used but is normally called a craniectomy. In March 2013, for the first time in the U. S., researchers replaced a large percentage of a patient's skull with a precision, 3D printed polymer implant. About 9 months later, the first complete cranium replacement with a 3D printed plastic insert was performed on a Dutch woman. She had been suffering from hyperostosis, which increased the thickness of her skull and compressed her brain. A study conducted in 2018 by the researchers of Harvard Medical School in Boston, funded by National Institutes of Health (NIH), suggested that instead of travelling via blood, there are "tiny channels" in the skull through which the immune cells combined with the bone marrow reach the areas of inflammation after an injury to the brain tissues.

Surgical alteration of sexually dimorphic skull features may be carried out as a part of facial feminization surgery or facial masculinization surgery, these reconstructive surgical procedures that can alter sexually dimorphic facial features to bring them closer in shape and size to facial features of the desired sex. These procedures can be an important part of the treatment of transgender people for gender dysphoria.

Artificial cranial deformation is a largely historical practice of some cultures. Cords and wooden boards would be used to apply pressure to an infant's skull and alter its shape, sometimes quite significantly. This procedure would begin just after birth and would be carried on for several years.

Like the face, the skull and teeth can also indicate a person's life history and origin. Forensic scientists and archaeologists use quantitative and qualitative traits to estimate what the bearer of the skull looked like. When a significant amount of bones are found, such as at Spitalfields in the UK and Jōmon shell mounds in Japan, osteologists can use traits, such as the proportions of length, height and width, to know the relationships of the population of the study with other living or extinct populations. The German physician Franz Joseph Gall in around 1800 formulated the theory of phrenology, which attempted to show that specific features of the skull are associated with certain personality traits or intellectual capabilities of its owner. His theory is now considered to be pseudoscientific.

In the mid nineteenth century, anthropologists found it crucial to distinguish between male and female skulls. An anthropologist of the time, James McGrigor Allan, argued that the female brain was similar to that of an animal. This allowed anthropologists to declare that women were in fact more emotional and less rational than men. McGrigor then concluded that women's brains were more analogous to infants, thus deeming them inferior at the time. However, later studies show that women's skulls are slightly thicker and thus men may be more susceptible to head injury than women. However, other studies shows that men's skulls are slightly thicker in certain areas. As well as some studies showing that females are more susceptible to head injury (concussion) than males. Men's skulls have also been shown to maintain density with age, which may aid in preventing head injury, while women's skull density slightly decreases with age. Male skulls can all have more prominent supraorbital ridges, glabella, and temporal lines. Female skulls generally have rounder orbits and narrower jaws. Male skulls on average have larger, broader palates, squarer orbits, larger mastoid processes, larger sinuses, and larger occipital condyles than those of females. Male mandibles typically have squarer chins and thicker, rougher muscle attachments than female mandibles.