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Branch of primatesthe branch of primates

Apes (collectively Hominoidea /h//ɒ//m//ᵻ/'/n//ɔɪ//d//i//.//ə/) are a clade of Old World simians native to sub Saharan Africa and Southeast Asia (though they were more widespread in Africa, most of Asia, and Europe in prehistory), which together with its sister group Cercopithecidae form the catarrhine clade, cladistically making them monkeys. Apes do not have tails due to a mutation of the TBXT gene. In traditional and non scientific use, the term ape can include tailless primates taxonomically considered Cercopithecidae (such as the Barbary ape and black ape), and is thus not equivalent to the scientific taxon Hominoidea. There are two extant branches of the superfamily Hominoidea: the gibbons, or lesser apes; and the hominids, or great apes. The family Hylobatidae, the lesser apes, include four genera and a total of 20 species of gibbon, including the lar gibbon and the siamang, all native to Asia. They are highly arboreal and bipedal on the ground. They have lighter bodies and smaller social groups than great apes. The family Hominidae (hominids), the great apes, include four genera comprising three extant species of orangutans and their subspecies, two extant species of gorillas and their subspecies, two extant species of panins (bonobos and chimpanzees) and their subspecies, and humans in a single extant subspecies.

Although the hominoid fossil record is still incomplete and fragmentary, there is now enough evidence to provide an outline of the evolutionary history of humans. Previously, the divergence between humans and other extant hominoids was thought to have occurred 15 to 20 million years ago, and several species of that time period, such as Ramapithecus, were once thought to be hominins and possible ancestors of humans. But, later fossil finds indicated that Ramapithecus was more closely related to the orangutan; and new biochemical evidence indicates that the last common ancestor of humans and non hominins (that is, the chimpanzees) occurred between 5 and 10 million years ago, and probably nearer the lower end of that range (more recent); see Chimpanzee–human last common ancestor (CHLCA).

Major studies of behaviour in the field were completed on the three better known "great apes", for example by Jane Goodall, Dian Fossey and Birutė Galdikas. These studies have shown that in their natural environments, the non human hominoids show sharply varying social structure: gibbons are monogamous, territorial pair bonders, orangutans are solitary, gorillas live in small troops with a single adult male leader, while chimpanzees live in larger troops with bonobos exhibiting promiscuous sexual behaviour. Their diets also vary; gorillas are foliovores, while the others are all primarily frugivores, although the common chimpanzee hunts for meat. Foraging behaviour is correspondingly variable. In November 2023, scientists reported, for the first time, evidence that groups of primates, including apes, and, particularly bonobos, are capable of cooperating with each other.

Apart from humans and gorillas, apes eat a predominantly frugivorous diet, mostly fruit, but supplemented with a variety of other foods. Gorillas are predominantly folivorous, eating mostly stalks, shoots, roots and leaves with some fruit and other foods. Non human apes usually eat a small amount of raw animal foods such as insects or eggs. In the case of humans, migration and the invention of hunting tools and cooking has led to an even wider variety of foods and diets, with many human diets including large amounts of cooked tubers (roots) or legumes. Other food production and processing methods including animal husbandry and industrial refining and processing have further changed human diets. Humans and other apes occasionally eat other primates. Some of these primates are now close to extinction with habitat loss being the underlying cause.

All the non human hominoids are generally thought of as highly intelligent, and scientific study has broadly confirmed that they perform very well on a wide range of cognitive tests—though there is relatively little data on gibbon cognition. The early studies by Wolfgang Köhler demonstrated exceptional problem solving abilities in chimpanzees, which Köhler attributed to insight. The use of tools has been repeatedly demonstrated; more recently, the manufacture of tools has been documented, both in the wild and in laboratory tests. Imitation is much more easily demonstrated in "great apes" than in other primate species. Almost all the studies in animal language acquisition have been done with "great apes", and though there is continuing dispute as to whether they demonstrate real language abilities, there is no doubt that they involve significant feats of learning. Chimpanzees in different parts of Africa have developed tools that are used in food acquisition, demonstrating a form of animal culture.